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<!DOCTYPE html>
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<head>
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  <title>Gilbert Lab | UCSF & Arc Institute</title>
  <meta name="description" content="CRISPR epigenome engineering, functional genomics, and cancer biology research at UCSF and Arc Institute">
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        <li><a href="#research" class="link-underline">Research</a></li>
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        <li><a href="#events" class="link-underline">Events</a></li>
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  <!-- Hero Section - Full viewport landing -->
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      <h1><span>Gilbert Lab</span></h1>
      <p class="hero-sub">Engineering the Epigenome</p>
      <div class="hero-affiliations">
        <a href="https://arcinstitute.org" target="_blank" class="link-underline">Arc Institute</a>
        <a href="https://www.ucsf.edu" target="_blank" class="link-underline">UCSF</a>
        <a href="https://cancer.ucsf.edu" target="_blank" class="link-underline">Helen Diller Cancer Center</a>
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  <!-- Research Section -->
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      <p class="section-label fade-in">Research</p>
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            <h3>Epigenome Engineering</h3>
          </div>
          <p>CRISPRi, CRISPRa, and CRISPRoff for programmable control of gene expression without cutting DNA.</p>
        </div>
        <div class="research-card">
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            <h3>Functional Genomics</h3>
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          <p>Genome-scale screens to map genetic interactions and identify synthetic lethal targets in cancer.</p>
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            <h3>Single-Cell / Perturb-seq</h3>
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          <p>High-throughput screens to map transcriptional responses across phenotypes and diverse cell types.</p>
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            <h3>Drug Resistance</h3>
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          <p>Modeling therapy response in prostate cancer, AML, glioblastoma, and lung cancer.</p>
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            <h3>COMET / Effector Discovery</h3>
          </div>
          <p>High-throughput screening of combinatorial dCas9 effector fusions to discover transcriptional modulators and map domain interactions.</p>
        </div>
        <div class="research-card">
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            <h3>Combinatorial Perturbations</h3>
          </div>
          <p>Engineered Cas12a for multiplexed chromatin editing and higher-order genetic interactions.</p>
        </div>
      </div>
    </div>
  </section>

  <!-- Team Section -->
  <section id="team">
    <div class="section-inner">
      <div class="team-header fade-in">
        <div>
          <p class="section-label">Team</p>
          <h2>Our People</h2>
        </div>
      </div>

      <!-- Principal Investigator -->
      <div class="pi-card fade-in">
        <img src="https://images.squarespace-cdn.com/content/v1/5bff2e82b40b9dd83d79f536/1543516234015-KJQ6G1X0RVWKKE3E99H6/image-asset.jpeg" alt="Luke Gilbert" class="pi-photo">
        <div class="pi-info">
          <h3>Luke Gilbert</h3>
          <p class="pi-title">Principal Investigator</p>
          <p class="pi-bio">Associate Professor at UCSF and Core Investigator at Arc Institute. PhD from MIT. Developed CRISPRi/a with Jonathan Weissman. NIH New Innovator. Pew-Stewart Scholar.</p>
          <div class="pi-links">
            <a href="mailto:luke@arcinstitute.org">Email</a>
            <a href="https://scholar.google.com/citations?user=wlqxB00AAAAJ" target="_blank">Scholar</a>
            <a href="https://profiles.ucsf.edu/luke.gilbert" target="_blank">UCSF</a>
          </div>
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      <!-- Scientists & Postdocs -->
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        <h4>Scientists & Postdocs</h4>
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            <div class="member-name">Gary Wang</div>
            <div class="member-role">Scientist</div>
          </div>
          <div class="member">
            <img src="https://images.squarespace-cdn.com/content/v1/5bff2e82b40b9dd83d79f536/744a75f2-0d78-4736-9dee-2aea70f69875/tempImageCKft7v.jpg" alt="Kewa Gao" class="member-photo">
            <div class="member-name">Kewa Gao</div>
            <div class="member-role">Scientist</div>
          </div>
          <div class="member">
            <img src="https://images.squarespace-cdn.com/content/v1/5bff2e82b40b9dd83d79f536/1626720776617-VEAASKECS1F3YG9LUQYB/IMG-3575.jpg" alt="Sarah Hsu" class="member-photo">
            <div class="member-name">Sarah Hsu</div>
            <div class="member-role">Fellow</div>
          </div>
          <div class="member">
            <img src="https://images.squarespace-cdn.com/content/v1/5bff2e82b40b9dd83d79f536/f6cb2e98-52a0-494f-a37f-d9225638887a/Tyler.jpg" alt="Tyler Fair" class="member-photo">
            <div class="member-name">Tyler Fair</div>
            <div class="member-role">Scientist</div>
          </div>
        </div>
      </div>

      <!-- Graduate Students -->
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        <h4>Graduate Students</h4>
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          <div class="member">
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            <div class="member-name">Ashir Borah</div>
            <div class="member-role">Graduate Student</div>
          </div>
          <div class="member">
            <img src="https://images.squarespace-cdn.com/content/v1/5bff2e82b40b9dd83d79f536/ec81d6c1-6ec8-42b2-9008-54f91a900b23/caroline.jpg" alt="Caroline Wilson" class="member-photo">
            <div class="member-name">Caroline Wilson</div>
            <div class="member-role">Graduate Student</div>
          </div>
        </div>
      </div>

      <!-- Staff -->
      <div class="members-section fade-in">
        <h4>Staff</h4>
        <div class="members-grid" id="staff-grid">
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            <div class="member-name">Ishwa Muzaffery</div>
            <div class="member-role">Lab Manager</div>
          </div>
          <div class="member">
            <img src="https://images.squarespace-cdn.com/content/v1/5bff2e82b40b9dd83d79f536/eae05cbe-4579-4c3e-a2d7-6e75d82602e5/IMG_1965.jpg" alt="Mandy Sun" class="member-photo">
            <div class="member-name">Mandy Sun</div>
            <div class="member-role">Lab Technician</div>
          </div>
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      </div>
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  </section>

  <!-- Publications Section -->
  <section id="publications">
    <div class="section-inner">
      <div class="pub-header fade-in">
        <div>
          <p class="section-label">Publications</p>
          <h2>Selected Work</h2>
        </div>
        <a href="https://www.ncbi.nlm.nih.gov/pubmed/?term=gilbert+la" target="_blank" class="link-underline">View all publications →</a>
      </div>

      <div class="pub-list" id="publications-list">
        <a href="https://doi.org/10.1038/s41587-025-02856-w" target="_blank" class="pub-item fade-in">
          <div class="pub-year">2025</div>
          <div>
            <div class="pub-title">Integrated epigenetic and genetic programming of primary human T cells</div>
            <div class="pub-meta">Goudy L et al. · <em>Nature Biotechnology</em></div>
          </div>
          <div class="pub-arrow">→</div>
        </a>
        <a href="https://doi.org/10.1038/s41588-025-02388-8" target="_blank" class="pub-item fade-in">
          <div class="pub-year">2025</div>
          <div>
            <div class="pub-title">PTGES3 as a direct modulator of androgen receptor function</div>
            <div class="pub-meta">Li H et al. · <em>Nature Genetics</em></div>
          </div>
          <div class="pub-arrow">→</div>
        </a>
        <a href="https://doi.org/10.1038/s41587-024-02224-0" target="_blank" class="pub-item fade-in">
          <div class="pub-year">2025</div>
          <div>
            <div class="pub-title">Engineered Cas12a for combinatorial chromatin perturbations</div>
            <div class="pub-meta">Hsiung CCS et al. · <em>Nature Biotechnology</em></div>
          </div>
          <div class="pub-arrow">→</div>
        </a>
        <a href="https://pubmed.ncbi.nlm.nih.gov/33838111/" target="_blank" class="pub-item fade-in">
          <div class="pub-year">2021</div>
          <div>
            <div class="pub-title">Genome-wide programmable transcriptional memory (CRISPRoff)</div>
            <div class="pub-meta">Nuñez JK et al. · <em>Cell</em></div>
          </div>
          <div class="pub-arrow">→</div>
        </a>
        <a href="https://pubmed.ncbi.nlm.nih.gov/25307932/" target="_blank" class="pub-item fade-in">
          <div class="pub-year">2014</div>
          <div>
            <div class="pub-title">Genome-scale CRISPR control of gene repression and activation</div>
            <div class="pub-meta">Gilbert LA et al. · <em>Cell</em></div>
          </div>
          <div class="pub-arrow">→</div>
        </a>
      </div>
    </div>
  </section>

  <!-- Lab Events Section - Photo Gallery -->
  <section id="events">
    <div class="section-inner">
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        <p class="section-label">Lab Life</p>
        <h2>Celebrations & Milestones</h2>
      </div>
      <div class="events-carousel fade-in" data-carousel>
        <button class="carousel-btn prev" type="button" aria-label="Previous events">←</button>
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            <article class="event-card event-slide">
              <img src="images/events/holiday-2025.jpg" alt="Holiday Lunch 2025">
              <div class="event-card-overlay">
                <div class="event-card-date">Dec 17, 2025</div>
                <div class="event-card-title">Holiday Lunch 2025</div>
                <div class="event-card-desc">& of course our annual white elephant</div>
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            </article>
            <article class="event-card event-slide">
              <img src="images/events/laine-graduation.jpg" alt="Laine's PhD Graduation">
              <div class="event-card-overlay">
                <div class="event-card-date">Nov 14, 2025</div>
                <div class="event-card-title">Laine's PhD Graduation</div>
                <div class="event-card-desc">Congratulations Dr. Goudy!</div>
              </div>
            </article>
            <article class="event-card event-slide">
              <img src="images/events/graduations-2025.jpg" alt="Graduations & Farewells">
              <div class="event-card-overlay">
                <div class="event-card-date">Mar 24, 2025</div>
                <div class="event-card-title">Graduations & Farewells</div>
                <div class="event-card-desc">Celebrating multiple milestones</div>
              </div>
            </article>
            <article class="event-card event-slide">
              <img src="images/events/lunar-2025.jpg" alt="Lunar New Year">
              <div class="event-card-overlay">
                <div class="event-card-date">Feb 7, 2025</div>
                <div class="event-card-title">Lunar New Year Happy Hour</div>
                <div class="event-card-desc">Gong hei fat choi!</div>
              </div>
            </article>
            <article class="event-card event-slide">
              <img src="images/events/white-elephant-2024.jpg" alt="White Elephant 2024">
              <div class="event-card-overlay">
                <div class="event-card-date">Dec 13, 2024</div>
                <div class="event-card-title">White Elephant 2024</div>
                <div class="event-card-desc">Annual holiday gift exchange</div>
              </div>
            </article>
            <article class="event-card event-slide">
              <img src="images/events/outing-2024.jpg" alt="Lab Outing 2024">
              <div class="event-card-overlay">
                <div class="event-card-date">Jul 18, 2024</div>
                <div class="event-card-title">Lab Outing 2024</div>
                <div class="event-card-desc">Summer lab gathering</div>
              </div>
            </article>
            <article class="event-card event-slide">
              <img src="images/events/ashir-quals.jpg" alt="Ashir Passed Quals">
              <div class="event-card-overlay">
                <div class="event-card-date">Mar 4, 2024</div>
                <div class="event-card-title">Ashir Passed Quals</div>
                <div class="event-card-desc">Congratulations Ashir!</div>
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              <img src="images/events/lunar-2024.jpg" alt="Lunar New Year 2024">
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                <div class="event-card-date">Feb 9, 2024</div>
                <div class="event-card-title">Lunar New Year 2024</div>
                <div class="event-card-desc">Happy Lunar New Year from Arc!</div>
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              <img src="images/events/white-elephant-2023.jpg" alt="White Elephant & Baby Shower">
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                <div class="event-card-date">Dec 8, 2023</div>
                <div class="event-card-title">White Elephant & Baby Shower</div>
                <div class="event-card-desc">Dual celebration for Sarah</div>
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              <img src="images/events/gokart-2023.jpg" alt="Go Kart Racing">
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                <div class="event-card-date">Nov 29, 2023</div>
                <div class="event-card-title">Go Kart Racing</div>
                <div class="event-card-desc">K1 Speed lab outing</div>
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            </article>
            <article class="event-card event-slide">
              <img src="images/events/candle-2023.jpg" alt="Candle Making">
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                <div class="event-card-date">Aug 25, 2023</div>
                <div class="event-card-title">Candle Making Happy Hour</div>
                <div class="event-card-desc">Creative lab social</div>
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            </article>
            <article class="event-card event-slide">
              <img src="images/events/amanda-exit.jpg" alt="Amanda's Exit Talk">
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                <div class="event-card-date">Aug 11, 2023</div>
                <div class="event-card-title">Amanda's Exit Talk</div>
                <div class="event-card-desc">Farewell and best wishes!</div>
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        <a href="protocols/amplification_protocol.pdf" class="protocol-card">
          <h3>CRISPR Library Re-amplification</h3>
          <p>Protocol for amplifying CRISPR libraries for screening experiments.</p>
          <span class="protocol-link">Download PDF →</span>
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        <a href="https://weissman.wi.mit.edu/crispr/" target="_blank" class="protocol-card">
          <h3>CRISPRa/i Screening Protocols</h3>
          <p>Comprehensive screening methodologies using CRISPR activation and interference approaches.</p>
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          <p>Code, tools, and computational resources from the Gilbert Lab.</p>
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        <h2>Where Are They Now</h2>
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          <div class="alumni-name">Aidan Winters</div>
          <div class="alumni-meta">Scientist II, Neptune Bio</div>
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        <div class="alumni-item">
          <div class="alumni-name">Alex Ge</div>
          <div class="alumni-meta">2017-2019 · Medical Student, UCSF</div>
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          <div class="alumni-name">Amanda Chung</div>
          <div class="alumni-meta">2019-2023 · Editas Medicine</div>
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          <div class="alumni-name">Becky Fu</div>
          <div class="alumni-meta">Scientist, Acrobat Genomics</div>
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          <div class="alumni-name">Ben Herken</div>
          <div class="alumni-meta">Scientist, Xiara Therapeutics</div>
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          <div class="alumni-name">Chris Hsiung</div>
          <div class="alumni-meta">Assistant Professor, UCSF</div>
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          <div class="alumni-name">Chris Yogodzinski</div>
          <div class="alumni-meta">2018-2020 · Graduate Student, UNC</div>
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          <div class="alumni-name">Garrett Wong</div>
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          <div class="alumni-name">Greg Pommier</div>
          <div class="alumni-meta">2020-2022 · Medical Student, Johns Hopkins</div>
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          <div class="alumni-name">Haolong Li</div>
          <div class="alumni-meta">Assistant Professor, Fred Hutch</div>
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          <div class="alumni-name">Jason Estep</div>
          <div class="alumni-meta">Senior Scientist, Terremoto Biosciences</div>
        </div>
        <div class="alumni-item">
          <div class="alumni-name">Jason Swindermann</div>
          <div class="alumni-meta">Medical School, MD/PhD, UCSF</div>
        </div>
        <div class="alumni-item">
          <div class="alumni-name">John Liu</div>
          <div class="alumni-meta">Assistant Professor, UCSF</div>
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        <div class="alumni-item">
          <div class="alumni-name">Laine Goudy</div>
          <div class="alumni-meta">Scientist, Xiara Therapeutics</div>
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        <div class="alumni-item">
          <div class="alumni-name">Lilly Scheibe</div>
          <div class="alumni-meta">2019-2021 · Harvard Medical School</div>
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        <div class="alumni-item">
          <div class="alumni-name">Nicholas Sambold</div>
          <div class="alumni-meta">Graduate Student, Stanford</div>
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        <div class="alumni-item">
          <div class="alumni-name">Raymond Dai</div>
          <div class="alumni-meta">Data Acquisition & Architecture, Eli Lilly and Company</div>
        </div>
        <div class="alumni-item">
          <div class="alumni-name">Sara Misiukiewicz</div>
          <div class="alumni-meta">2020-2022 · Graduate Student, UCSF</div>
        </div>
        <div class="alumni-item">
          <div class="alumni-name">Shaheen Kabir</div>
          <div class="alumni-meta">2018-2021 · Graphite Bio</div>
        </div>
        <div class="alumni-item">
          <div class="alumni-name">Tom O'Loughlin</div>
          <div class="alumni-meta">2019-2022 · Assistant Professor, Mount Sinai</div>
        </div>
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          <strong>Location</strong><br>
          Arc Institute<br>
          3181 Porter Drive<br>
          Palo Alto, CA 94304
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          <a href="mailto:luke@arcinstitute.org" class="link-underline">luke [at] arcinstitute.org</a>
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          <a href="https://arcinstitute.org" target="_blank" class="link-underline">Arc Institute</a><br>
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